On Zanifar, Ratchet uses a time portal created by Sigmund to travel back in time two years earlier and rescue Orvus, who is being tortured by Nefarious. The Lombaxes find the Eye and use it to communicate with Clank, who brings them up to speed about his father's plight and implores them to go to Zanifar to save him.
Whilst searching for the Eye on Lumos, Alister hesistantly reveals that he was indirectly responsible for the near-extinction of the Lombaxes but he plans to change that by using the Clock to rewrite history. Meanwhile, Clank learns about the Clock and eavesdrops on Sigmund, indirectly learning that the real Orvus disappeared two years ago.
The Lombaxes team up and head to Terachnos to uncover the location of the Obsidian Eye, which allows the user to communicate with the inhabitants of the Great Clock, specifically Clank. On Torren IV, Ratchet is on the hunt for Alister, who is also a Lombax after initially mistaking Ratchet for an assassin and recognizes him as the son of Kaden. Sigmund helps Clank learn more about the Clock, with the help of his father Orvus in his subconsciousness, unaware that they are being spied on by Lawrence under Nefarious' orders. Clank is revived by Sigmund, the Junior Caretaker of the Great Clock, a machine that keeps time together. With his ship repaired by the Zoni, Ratchet journeys to Vorselon's warship and rescues the hostages. Lord Vorselon, Nefarious' personal assassin, launches an attack on the planet in search of someone named "Alister Azimuth" and captures both the Fongoids and Qwark for more information. Meanwhile in the Phylax Sector, Ratchet and Captain Qwark are struck by a time anomaly and crash-land on the planet Quantos in their search for Clank. Nefarious tells Lawrence to spare Clank, saying that he has more for him in mind. In retaliation, Nefarious attempts to kill them, inadvertently allowing Clank to escape, only for the latter to be immobilized by Lawrence moments later. Nefarious has been assigned by the Zoni to fix Clank, but in reality, he is after a room called the "Orvus Chamber", to which the Zoni deny him access. Ratchet and Clank A Crack In Time is based after the events of Ratchet and Clank Tools of Destruction and Ratchet and Clank Quest for Booty.
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